Andy Gardner is an evolutionary biologist at the University of St Andrews. His work lies at the interface of theoretical population genetics and behavioural ecology, and focuses on social evolution. Andy is fascinated by Darwinian adaptation: natural selection expains the appearance of design in the living world, but there remains controversy as to whether this design manifests at the level of genes, individuals or whole societies. Social evolution provides a window onto this problem, by pitting the interests of genes, individuals and societies against each other. Andy develops general theory and mathematical models tailored to the biology of specific species, from microbes to insects to humans.